Fst
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Fst is a measure of population divergence. It measures the reduction in heterozygosity in a subdivided population compared to that in large, random mating population with the same allele frequencies.
Formulae
References
Nielsen R, Mountain JL, Huelsenbeck JP, Slatkin M (1998) Maximum-Likelihood Estimation of Population Divergence Times and Population Phylogeny in Models Without Mutation. Evolution 52:669-677